rugae (n.)youdaoicibaDictYouDict[rugae 词源字典]
plural of ruga (1775), from Latin ruga "a wrinkle in the face," from PIE *rug-, variant of *reue- (2) "to smash, knock down, tear up, dig up, uproot" (see rough (adj.)).[rugae etymology, rugae origin, 英语词源]