pronogradeyoudaoicibaDictYouDict[pronograde 词源字典]
"Of a primate: holding the body more or less horizontal; (of posture) not erect; quadrupedal. Contrasted with orthograde", Early 20th cent.; earliest use found in Arthur Keith (1866–1955), museum curator and palaeoanthropologist. From classical Latin prōnus prone + -o- + -grade.[pronograde etymology, pronograde origin, 英语词源]